Speculation-
Engines - more thrust?
Airframe - hardened and strengthened to withstand greater G forces for better maneuverability?
Communications gear to survive any type of EMP/nuclear event?
Advanced radar and potentially radar jamming gear to confuse missiles and enemy aircraft?
Weapons systems - air-to-air and/or air-to-ground missiles?
Obviously a nice custom interior with its own security features to protect the president.
I’m just guessing - but something, or multiple things, have to account for an extraordinary price.